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activity adenosine adenylate adsorbed alanine algae amino acids aqueous solution atmosphere atoms bacteria Biochem Biol biological carbon carboxylation catalyst Chem chemical evolution chloroplast chromatogram chromatography clay coacervate CODE complex compounds concentration condensation cyanide cytochrome DAYHOFF degassing electron energy enzyme ethanethiol eukaryotic Evol evolutionary EVREINOVA experiments fatty acids formaldehyde formation formed formose reaction fraction function GC-content genetic glycine hydrogen hydrolysis interaction interstellar irradiation matrix membrane metabolic meteorites mixture molecular molecules montmorillonite NaCl nitriles nucleic acids nucleotide obtained oligomers OPARIN organic ORGEL origin oxidation oxygen peak peptide phosphate photosynthetic planet poly polymerization polymers polynucleotides PONNAMPERUMA prebiotic Precambrian precursors present primitive earth prokaryotic protein proteinoid radiation ratio reaction redox reduced samples sequence shown in Fig structure surface synthesis Table temperature thymine tion TOUPANCE tRNA type cells uracil values yield