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by selecting such slight differences, yet they ignore all general arguments, and refuse to sum up in their minds slight differences accumulated during many successive generations. LMay not those naturalists who, knowing far less of the laws of inheritance than does the breeder, and knowing no more than he does of the intermediate links in the long lines of descent, yet admit that many of our domestic races have descended from the same parents—may they not learn a lesson of caution, when they deride the idea of species in a tate of nature being lineal descendants of other species ‘I

Selection—Let us now briefly consider the steps by which domestic races have been produced, either from one or from several allied species. Some little effth may be attributed to the direct actio'rT-of theirtgrnal mm enema some little to habit I'mhe would be aTBold“m‘aii“who“wouId‘acc0unt by'such agencies for the differences of a dray and race horse, a greyhound and bloodhound, a carrier and tumbler pigeon. (One of the most remarkable features in our domesticated races is that we see in them adaptation, not indeed to the animal’s or plant’s own good, but to man’s use or fancyfj Some variations useful to him have probably arisen suddenly, or by one step; many botanists, for instance, believe that the fuller’s teazle, with its hooks, which cannot be rivalled by any mechanical contrivance, is only a variety of the wild Dipsacus; and this amount of change may have suddenly arisen in a seedling. So it has probably been with the turnspit dog; and this is known to have been the case with the ancon sheep. But when we compare the dray-horse and race-horse, the dromedary and camel, the various breeds of sheep fitted either for/cultivated land or mountain pasture, with the wool of "‘one breed good for one purpose, and that of another breed for another purpose; when we compare the many breeds of dogs, each good for man in very different ways; when we compare the game-cock, so pertinacious in battle, with other breeds so little quarrelsome, with “ everlasting layers” which never desire to sit, and with the bantam so small and elegant; when we compare the host of agricultural, culinary, orchard, and flower-garden races of plants, most useful to man at different seasons and for different purposes, or so beautiful in his eyes, we must, think, look further than to mere variabilityfi We cannot. suppose that all __the breeds

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38 VARIATION UNDER DOMESTICATION

were suddenly-produced as perfectland as useful aslwe muee them; indeed, in several cases, we know that this has not been their history. The key is man’s power of accumulative selection: nature gives successive varia-_ tions; man adds them up in certain directions useful to him. In this sense he may b said to make for himself useful breeds.) Mb»- Lai‘ Mfr» hismwérmuw ‘ .

The great power of this principle of selection is not hypothetical. It is certain that several of our eminent breeders have, even within a single lifetime, modified to a large extent some breeds of cattle and sheep. In order fully to realise what they have done, it is almost necessary to read several of the many treatises devoted to this subject, and to inspect the animals. Breeders habitually speak of an animal’s organisation as something quite plastic, which they can model almost as they please. If I had space I could quote numerous passages to this. effect from highly competent authorities. Youatt, who was probably better acquainted with the works of agriculturists than almost any other individual, and who was himself a very good judge of an animal, speaks of the principle of selection as “that which enables the agriculturist not only to modify the character of his flock, but to change it altogether. It is the magician’s wand, by means of which he may summon into life whatever form and mould he pleases." Lord Somerville, speaking of what breeders have done for sheep, says ;— “ It would seem as if they had chalked out upon a wall a form perfect in itself, and then had given it existence.” That most skilful breeder, Sir John Sebright, used to say, with respect to pigeons, that “ he would produce any given feather in three years, but it would take him six years to obtain head and beak.” In Saxony the importance of the principle of selection in regard to merino sheep is so fully recognised, that men follow it as a trade: the sheep are placed on a table and are studied, like a picture by a connoisseur; this is done three times at intervals of months, and the sheep are each time marked and classed, so that the very best may ultimately be selected for breeding.

What English breeders have actually effected is proved by the enormous prices given for animals with a good pedigree; and these have now been exported to almost every quarter of the world. L'll‘he improvement is by no means generally due to‘ crossing different breeds; all the best breeders are strongly opposed to this practice, except sometimes amongst closely allied sub-breeds. And when a cross has been made, the closest selection is far more indispensable even than in ordinary cases. If selection consisted merely in separating some very distinct variety, and breeding from it, the principle would be so obvious as hardly to be worth notice; but its importance consists in the great effect produced by the accumulation in one direction, during successive generations, of differences absolutely inappreciable by an uneducated eye—differen_ es which I for one have vainly attempted to appreciate. Not one man in a thousand has accuracy of eye and judgment sufficient to become an eminent breeder. If gifted with these qualities, and he studies his subject for years, and devotes his lifetime to it with indomitable perseverance, he will succeed, and may make great improvements; if he wants any of these qualities, he will assuredly fail. Few would readily believe in the natural capacity and years of practice requisite to become even a skilful pigeon-fancier.

The same principles are followed by horticulturists; but the variations are here often more abrupt. No one supposes that our choicest productions have been produced by a single variation from the aboriginal stock. We have proofs that this is not so in some cases in which exact records have been kept; thus, to give a very trifling instance, the steadily-increasing size of the common gooseberry may be quoted. We see an astonishing improvement in many florists’ flowers, when the flowers of the present day are compared with drawings made only twenty or thirty years ago. When a race of plants is once pretty well established, the seed-raisers do not pick out the best plants, but merely go over their seedbeds, and pull up the “ rogues,” as they call the plants that deviate from the proper standard. With animals this kind of selection is, in fact, also followed; for hardly any one is so careless as to allow his worst animals to breed.

In regard to plants, there is another means of observing the accumulated effects of selection—namely, by com— paring the diversity of flowers in the different varieties of the same species in the flower-garden ; Y the diversity of leaves, pods, or tubers, or whatever part is valued, in the kitchen-garden, in comparison with the flowers of the same varieties; and the diversity of fruit of the same

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species in the orchard, in comparison With the leaves and flowers of the same set of varieties. See how different the leaves of the cabbage are, and how extremely alike the flowers ; how unlike the flowers of the heartsease are, and how alike the leaves; how much the fruit of the different kinds of gooseberries differ in size, colour, shape, and hairiness, and yet the flowers present very slight differences. It is not that the varieties which differ largely in‘ some one point do not differ at all in other points; this is hardly ever, I speak after careful observation, perhaps never, the case. LThe laws of correlation of growth, the importance of which should never be overlooked, will ensure some differences ; but, as a general rule, I cannot doubt that the continued selection of slight variations, either in the leaves, the flowers, or the fruit, will produce races differing from each other chiefly in these characters.) QM“ I “Mined-l» VM-d-e

< It may be objected that the principle of selection has been reduced to methodical practice for scarcely more than three-quarters of a century it has certainly been more attended to of late years, and many treatises have been published on the subject; and the result has been, in a corresponding degree, rapid and important. [ But it is very far from true that the principle is a modern discovery] I could give several references to the full acknowledgment of the importance of the principle in works of high antiquity. In rude and barbarous periods of English history choice animals were often imported, and laws were passed to prevent their exportation: the destruction of horses under a certain size was ordered, and this may be compared to the “ roguing ” of plants by nurserymen. The principle of selection I find distinctly given in an ancient Chinese encyclopaedia. Explicit rules are laid down by some of the Roman classical writers. From passages in Genesis, it is clear that the colour of domestic animals was at that early period attended to. Savages now sometimes cross their dogs with wild canine animals, to improve the breed, and they formerly did so, as is attested by passages in Pliny. The saVages in South Africa match their draught cattle by colour, as do some of the Esquimaux their teams of dogs. Livingstone shows how much good domestic breeds are valued by the negroes of the interior of Africa who have not associated with Europeans. Some of these facts do not show actual selection, but they show that the breeding

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of domestic animals was carefully attended to in ancient times, and is now attended to by the lowest savages. {It would, indeed, have been a strange fact, had attention not been paid to breeding, for the inheritance of good and bad qualities is so obvious;

At the present time, eminent breeders try by methodical selection, with a distinct object in view, to make a new strain or sub-breed, superior to anything existing in the country. But, for our purpose, a kind of Selection, which may be called Unconscious, and which results from every one trying to possess and bree from the best individual animals, is more important. Thus, a man who intends keeping pointers naturally tries to get as good dogs as he can, and afterwards breeds from his own best dogs, but he has no wish or expectation of permanently altering the breed?) Nelr_e_rthelgss__vve_§i/ziy infer that ___this process, conlinuiiduring centuries, wogld imflprove and modify any b31291, in the same way as' Bakewell, Collins, 250., By this very same process, only carried on more methodically, did greatly modify, even during their own lifetimes, the forms and qualities of their cattle. Slow and insensible changes of this kind could never be recognised unless actual measure; ments or careful drawings of the breeds in question had been made long ago, which might serve for comparison. In some cases, however, unchanged, or but little changed individuals of the same breed may be found in less civilised districts, where the breed has been less improved. There is reason to believe that King Charles’s spaniel has been unconsciously modified to a large extent since the time of that monarch.ZSome highly competent authorities are convinced that the setter is directly derived from the1 spaniel, and has probably been slowly altered from it. Q It is known that the English pointer has been greatly changed within the last century, and in this case the change has, it is believed been chiefly effected by crosses with the fox-hound; but what concerns us is, that the change has been effected unconsciously and gradually, Jand yet so effectually, that, though the old Spanish pointer certainly came from Spain, Mr. Borrow has not seen, as I am informed by him, any native dog in Spain like our pointer.

By a similar process of selection, and by careful training, the whole body of English racehorses have come to surpass in fleetness and size the parent Arab stock, so

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