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as varieties, but which have all been ranked by botanists as species; and in making this list he has omitted

many trifling varieties, but which nevertheless have been ranked by some botanists as species, and he has entirely omitted several highly polymorphic genera. Under genera, including the most polymorphic forms, Mr. Babington gives 251 species, whereas Mr. Bentham gives only 112,-a difference of 139 doubtful forms! Amongst animals which unite for each birth, and which are highly locomotive, doubtful forms, ranked by one zoologist as a species and by another as a variety, can rarely be found within the same country, but are common in separated areas. How many of the birds and insects in North America and Europe, which differ very slightly from each other, have been ranked by one eminent naturalist as undoubted species, and by another as varieties, or, as they are often called geographical races! Mr. Wallace, in several valuable papers on the various animals, especially on the Lepidoptera, inhabiting the islands of the great Malayan archipelago, shows that they may be classed under four heads, namely, as variable forms, as local forms, as geographical races or sub-species, and as true representative species. The first or variable forms vary much within the limits of the same island. The local forms are moderately constant and distinct in each separate island; but when all from the several islands are compared together, the differences are seen to be so slight and graduated, that it is impossible to define or describe them, though at the same time the extreme forms are sufficiently distinct. The geographical races or sub-species are local forms completely fixed and isolated; but as they do not differ from each other by

strongly marked and important characters, “there is no
possible test but individual opinion to determine which
of them shall be considered as species and which as
varieties.” Lastly, representative species fill the same
place in the natural economy of each island as do the
local forms and sub-species; but as they are distin-
guished from each other by a greater amount of differ-
ence than that between the local forms and sub-species,
they are almost universally ranked by naturalists as
true species. Nevertheless, no certain criterion can
possibly be given by which variable forms, local
forms, sub-species, and representative species can be
recognised.
Many years ago, when comparing, and seeing others
compare, the birds from the closely neighbouring islands
of the Galapagos archipelago, one with another, and
with those from the American mainland, I was much
struck how entirely vague and arbitrary is the dis-
tinction between species and varieties. On the islets of
the little Madeira group there are many insects which
are characterised as varieties in Mr. Wollaston's ad-
mirable work, but which would certainly be ranked as
distinct species by many entomologists. Even Ireland
has a few animals, now generally regarded as varieties,
but which have been ranked as species by some zo-
ologists. Several experienced ornithologists consider
our British red grouse as only a strongly-marked race
of a Norwegian species, whereas the greater number
rank it as an undoubted species peculiar to Great
Britain. A wide distance between the homes of two
doubtful forms leads many naturalists to rank them
as distinct species; but what distance, it has been well
asked, will suffice; if that between America and Europe

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is ample, will that between Europe and the Azores, or Madeira, or the Canaries, or between the several islets of these small archipelagos, be sufficient? Mr. B. D. Walsh, a distinguished entomologist of the United States, has described what he calls Phytophagic varieties and Phytophagic species. Most vegetablefeeding insects live on one kind of plant or on one group of plants; some feed indiscriminately on many kinds, but do not in consequence vary. In several cases, however, insects found living on different plants, have been observed by Mr. Walsh to present in their larval or mature state, or in both states, slight, though constant differences in colour, size, or in the nature of their secretions. In some instances the males alone, in other instances both males and females, have been observed thus to differ in a slight degree. When the differences are rather more strongly marked, and when both sexes and all ages are affected, the forms are ranked by all entomologists as good species. But no observer can determine for another, even if he can do so for himself, which of these Phytophagic forms ought to be called species and which varieties. Mr. Walsh ranks the forms which it may be supposed would freely intercross, as varieties; and those which appear to have lost this power, as species. As the differences depend on the insects having long fed on distinct plants, it cannot be expected that intermediate links connecting the several forms should now be found. The naturalist thus loses his best guide in determining whether to rank doubtful forms as varieties or species. This likewise necessarily occurs with closely allied organisms, which inhabit distinct continents or islands. When, on the other hand, an animal or plant ranges over the same continent, or inhabits many islands in the same archipelago, and presents different forms in the different areas, there is always a good chance that intermediate forms will be discovered which will link together the extreme states, and these are then degraded to the rank of varieties. Some few naturalists maintain that animals never present varieties; but then these same naturalists rank the slightest difference as of specific value; and when the same identical form is met with in two distant countries, or in two geological formations, they believe that two distinct species are hidden under the same dress. The term species thus comes to be a mere useless abstraction, implying and assuming a separate act of creation. It is certain that many forms, considered by highly-competent judges to be varieties, resemble species so completely in character, that they have been thus ranked by other highly-competent judges. But to discuss whether they ought to be called species or varieties, before any definition of these terms has been generally accepted, is vainly to beat the air. Many of the cases of strongly-marked varieties or doubtful species well deserve consideration; for several interesting lines of argument, from geographical distribution, analogical variation, hybridism, &c., have been brought to bear in the attempt to determine their rank; but space does not here permit me to discuss them. Close investigation, in many cases, will no doubt bring naturalists to agree how to rank doubtful forms. Yet it must be confessed that it is in the best known countries that we find the greatest number of them. I have been struck with the fact, that if any animal or plant in a state of nature be highly useful to man, or from any cause closely attracts his attention, varieties of it will almost universally be found recorded. These varieties, moreover, will often be ranked by some authors as species. Look at the common oak, how closely it has been studied; yet a German author makes more than a dozen species out of forms, which are almost universally considered by other botanists to be varieties; and in this country the highest botanical authorities and practical men can be quoted to show that the sessile and pedunculated oaks are either good and distinct species or mere varieties. I may here allude to a remarkable memoir lately published by A. de Candolle, on the oaks of the whole world. No one ever had more ample materials for the discrimination of the species, or could have worked on them with more zeal and sagacity. He first gives in detail all the many points of structure which vary in the several species, and estimates numerically the relative frequency of the variations. He specifies above a dozen characters which may be found varying even on the same branch, sometimes according to age or development, sometimes without any assignable reason. Such characters are not of course of specific value, but they are, as Asa Gray has remarked in commenting on this memoir, such as generally enter into specific definitions. De Candolle then goes on to say that he gives the rank of species to the forms that differ by characters never varying on the same tree, and never found connected by intermediate states. After this discussion, the result of so much labour, he emphatically remarks: “They are mistaken, who repeat that the greater part of our species are clearly

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